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The nucleus contains the cell’s genome. It is bounded by a
membrane that consists of a pair of unit membranes separated by a space of variable thickness. The inner membrane is
usually a simple sac, but the outermost membrane is, in many
places, continuous with the endoplasmic reticulum (ER). The
nuclear membrane exhibits selective permeability because of
pores, which consist of a complex of several proteins whose
function is to import substances into and export substances
out of the nucleus. The chromosomes of eukaryotic cells contain linear DNA macromolecules arranged as a double helix.
They are only visible with a light microscope when the cell is
undergoing division and the DNA is in a highly condensed
form; at other times, the chromosomes are not condensed
and appear as in Figure 2-2. Eukaryotic DNA macromolecules are associated with basic proteins called histones that
bind to the DNA by ionic interactions.
A structure often visible within the nucleus is the nucleolus, an area rich in RNA that is the site of ribosomal RNA
synthesis (see Figure 2-2). Ribosomal proteins synthesized in
the cytoplasm are transported into the nucleolus and combine with ribosomal RNA to form the small and large subunits of the eukaryotic ribosome. These are then exported to
the cytoplasm, where they associate to form an intact ribosome that can function in protein synthesis
The nucleus contains the cell’s genome. It is bounded by a
membrane that consists of a pair of unit membranes separated by a space of variable thickness. The inner membrane is
usually a simple sac, but the outermost membrane is, in many
places, continuous with the endoplasmic reticulum (ER). The
nuclear membrane exhibits selective permeability because of
pores, which consist of a complex of several proteins whose
function is to import substances into and export substances
out of the nucleus. The chromosomes of eukaryotic cells contain linear DNA macromolecules arranged as a double helix.
They are only visible with a light microscope when the cell is
undergoing division and the DNA is in a highly condensed
form; at other times, the chromosomes are not condensed
and appear as in Figure 2-2. Eukaryotic DNA macromolecules are associated with basic proteins called histones that
bind to the DNA by ionic interactions.
A structure often visible within the nucleus is the nucleolus, an area rich in RNA that is the site of ribosomal RNA
synthesis (see Figure 2-2). Ribosomal proteins synthesized in
the cytoplasm are transported into the nucleolus and combine with ribosomal RNA to form the small and large subunits of the eukaryotic ribosome. These are then exported to
the cytoplasm, where they associate to form an intact ribosome that can function in protein synthesis
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